Scott Cummings and Kyle Steyn make Scotland squad

Gregor Townsend left two spaces open - for a second-rower and a winger - when he named a 42-man squad three weeks ago.

And 22-year-old lock Cummings and versatile back Steyn, who can play at centre or out wide, have won the race after a strong finish to the season, which culminated in a narrow Guinness PRO14 final defeat by Leinster on Saturday.

That means no spot for Gray, who won the last of his 65 caps against Italy last year before back and hip injuries ruled him out for the best part of a year.

The former Glasgow player is back to full fitness but has only made six starts for Toulouse this season and has not featured enough in recent weeks to force his way into Townsend's plans.

Townsend said: "We are fortunate to have a number of very good second-rows available to us right now. With Richie Gray, Tim Swinson and Andrew Davidson also in contention for this remaining place in our World Cup squad, Scott's selection is a testament to how well he has performed throughout the season and in particular over the past few weeks.

"Scott has played very well in some crucial games for Glasgow and has really taken his game to the opposition. He was impressive in his ball carrying, defence and his all-round work-rate on Saturday against Leinster and we look forward to working with him and the other four second-rows in our squad."

Swinson has been troubled by an ankle injury recently, while Davidson has not played much for Newcastle ahead of his return to Glasgow.

South Africa-born Steyn, whose mother is from Glasgow, has forced his way in after some impressive performances for Scotland Sevens and Warriors this season, following his move to Scotland.

The 25-year-old only joined Warriors in February to integrate ahead of a one-year contract for next season, but his rise has been rapid.

Townsend said: "Kyle has made an excellent impact since his debut for Glasgow earlier this season and has displayed strong defensive capabilities as well as a providing go-forward for his team when carrying ball.

"His versatility is a bonus. He started out playing on the wing for Glasgow, has moved into the centre and has performed equally well in both roles.

"With every game he has looked more assured and, from our point of view, he will primarily be considered a winger who can cover the centre position.

"We have others in our squad that we view more as centre/wings but I'm sure over the course of the summer Kyle will get the opportunity to show what he can do in both positions."